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This course provides a comprehensive introduction to basic principles of macromolecular structure, stability, and interactions. Major topics include protein structure; protein thermodynamics and folding; nucleic acid structure and stability; principles of intermolecular recognition; and principles and practice of ligand binding analysis. Special emphasis is placed on understanding, in macromolecular systems, the relationships between structure and stability; the molecular origins of cooperative effects; and the relationships between covalent and non-covalent properties.